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However, perforation of the tympanic membranes severely reduced sensitivity to low frequencies, indicating that sensitivity to low frequencies is mediated primarily by the ears and cannot be attributed to some other sensory modality.In the terrestrial slugs Limax, most of the photosensory information is thought to be acquired by an eye located on the superior tentacles, by which the slugs avoid light. Recent studies, however, suggested that the brain also plays a role as a photosensor in their negative phototaxis behavior. In the prese