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MYB-bHLH-WDR transcription factor (TF) complexes (MBWs) regulate Arabidopsis seed coat development including mucilage and tannin biosynthesis. The R2R3 MYBs MYB5, MYB23 and TRANSPARENT TESTA2 (TT2) participate in the MBW complexes with the WD-repeat protein TRANSPARENT TESTA GLABRA1 (TTG1). These complexes regulate GLABRA2 (GL2) and TTG2 expression in developing seeds. Microarray transcriptome analysis of ttg1-1 and wild-type (Ler) developing seeds identified 246 TTG1-regulated genes which include all known metabolic genes of the tannin

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