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These di- and tricarboxylates are imported into the cell because of the divalent anion sodium symporter (DASS) category of plasma membrane transporters, which contains both cotransporters and exchangers. While DASS proteins transport substrates via an elevator apparatus, to date structures are merely designed for just one DASS cotransporter protein in a substrate-bound, inward-facing state. We report numerous cryo-EM and X-ray structures in four various says, including three hitherto unseen state
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