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Orienting to gaze-direction is widespread among animal species, but evidence for spontaneous use of gesture for direction is limited [1]. Remarkably, African elephants (Loxodonta africana) have been found able to follow human pointing, including subtle actions in which the contralateral hand is used, and in which the body silhouette is not broken [2,3]. The natural origin of this ability is puzzling, as the species is not reported to use trunk- or limb-gesture for showing directions [4]. One natural gesture, the 'periscope-sniff' presu
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