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© Prof. H.S. Srivastava Foundation for Science and Society 2020.Heatwaves are becoming much more frequent with environment warming and that can impact tree development and reproduction. Eucalyptus parramattensis can cope with an extreme heatwave in the field via transpiratory air conditioning and enhanced leaf thermal tolerance that protected foliar tissues from photo-inhibition and photo-oxidation during natural midday irradiance. Here, we explored whether alterations in fo