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Expected final online publication date for the Annual Review of Genomics and Human Genetics Volume 22 is August 2021. Please see http//www.annualreviews.org/page/journal/pubdates for revised estimates.The gametophyte represents the sexual phase in the alternation of generations in plants; the other, nonsexual phase is the sporophyte. Here, we review the evolutionary origins of the male gametophyte among land plants and, in particular, its ontogenesis in flowering plants. The highly reduced male gametophyte of angiosperm plants i
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