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The leading-edge vortex grows slower and stays close-bound to the wing throughout the majority of the stroke-cycle. This requires less aerodynamic power and increases the hovering efficiency by 93% but sacrifices 43% of the maximum lift in the process. In all cases, a leading-edge vortex is fed by vorticity through the leading edge shear-layer which makes the shear-layer velocity a good indicator for the growth of the vortex and its impact on the aerodynamic forces. We estimate the shear-layer velocity at the leading edge solely from the
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