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For permissions, please email journals.permissions@oup.com.Convergent evolution is pervasive in nature, but it is poorly understood how various constraints and natural selection limit the diversity of evolvable phenotypes. Here, we analyze the transcriptome across fruiting body development to understand the independent evolution of complex multicellularity in the two largest clades of fungi-the Agarico- and Pezizomycotina. Despite 650 million years of divergence between these clades, we find that very similar sets of genes have conv
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