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We report a number of negative carry-over effects of non-breeding activity on breeding, which were generally stronger in shyer individuals more active winters were followed by later breeding phenology and poorer breeding performance in shy birds, but these effects were weaker or undetected in bolder individuals. Our study quantifies individual variability in the strength of carry-over effects on breeding and provides a mechanism explaining widespread differences in individual reproductive success.The capacity for parents to influence of
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