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Expected final online publication date for the Annual Review of Entomology, Volume 66 is January 11, 2020. Please see http//www.annualreviews.org/page/journal/pubdates for revised estimates.Distant and predictable features in the environment make ideal compass cues to allow movement along a straight path. Ball-rolling dung beetles use a wide range of different signals in the day or night sky to steer themselves along a fixed bearing. These include the sun, the Milky Way, and the polarization pattern generated by the moon. Almost two decades of res