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Here we report the finding of elaborate age polyethism in a social aphid (Hemiptera). Tuberaphis styraci is a gall-forming aphid in which monomorphic very first instar nymphs differentiate into regular nymphs and soldiers upon second instar molt. Soldiers neither grow nor reproduce but perform gall cleaning and colony protection. Utilizing an artificial diet rearing system, we obtained age-defined groups of soldiers and monitored their particular personal