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The ultrametric method differs from the traditional autodiametric method in that PVOs can be separated by ultrasonication and then enumerated, and ovarian homogeneity is not a mandatory requirement per se. The method is designed for determinate spawners but might be extended to include indeterminate spawners.An amendment to this paper has been published and can be accessed via a link at the top of the paper.Survival in complex environments necessitates a flexible navigation system that incorporates memory of recent behavior and association