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Our results were noticeably consistent across soil typologies, both for fruitbodies growing singly and in clusters, indicating that early-developing fruitbody characters did not influence characters linked to subsequent morphogenetic stages. The lack of links among stages opens new perspectives for pre-harvest quality management with stage-specific cultivation practices.Here, we aimed to boost antibacterial performance of cellulose fibers for paper sheet application. Therefore, TiO2 nanoparticles have been used with controlled loading