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eCO and N enhanced both g and A , but to differing degrees; g was more strongly influenced by N through increases in stomatal density while A was more strongly stimulated by eCO . There was a positive relationship between g and A that was not modified by eCO or N , individually or in combination. Changes in stomatal features co-occur with previously described responses of P. australis to eCO and N . Complementary responses of stomatal length and density to these global change factors may facilitate greater stomatal conductance and carb