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Results indicate that a reduction of the habitat suitability of the lung lichen across Italy should be expected in the face of climate change and that this is coupled with a loss of suitable substrate. This situation seems to be determined by two main processes that act simultaneously 1) a partial reduction of the spatial overlap between the climatic niche of the lung lichen and that of its host tree species, and 2) the invasion of native woods by black locust. The case of lung lichen and black locust in Italy highlights that epiphytes