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All rights reserved. For permissions, please email journals.permissions@oup.com.Plant nitrogen (N) acquired mainly in forms of nitrate and ammonium from soil dominates growth and development, and high-yield crop production relies heavily on N fertilization. The mechanisms of root adaptation to altered supply of N forms and concentrations have been well characterized and reviewed, while reports concerning effects of N on the architecture of vegetative and reproductive organs are limited and are widely dispersed in the literature. In this